The experience of the breeders was quite inadequate to the use which
Darwin made of it. It was neither scientific, nor critically accurate.
Laws of variation were barely conjectured; the different types of
variability were only imperfectly distinguished. The breeders'
conception was fairly sufficient for practical purposes, but science
needed a clear understanding of the [6] factors in the general process
of variation. Repeatedly Darwin tried to formulate these causes, but the
evidence available did not meet his requirements.
Quetelet's law of variation had not yet been published. Mendel's claim
of hereditary units for the explanation of certain laws of hybrids
discovered by him, was not yet made. The clear distinction between
spontaneous and sudden changes, as compared with the ever-present
fluctuating variations, is only of late coming into recognition by
agriculturists. Innumerable minor points which go to elucidate the
breeders' experience, and with which we are now quite familiar, were
unknown in Darwin's time. No wonder that he made mistakes, and laid
stress on modes of descent, which have since been proved to be of minor
importance or even of doubtful validity.
Notwithstanding all these apparently unsurmountable difficulties, Darwin
discovered the great principle which rules the evolution of organisms.
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