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Vries, Hugo de, 1848-1935

"Species and Varieties, Their Origin by Mutation"

The first is the size of the cultures required for the
resulting amelioration. In other words, would it have been possible to
attain an average of 20 rows in a single experiment? This is a matter of
calculation, and the calculation must be based upon the experience
related above, that the progression in the case of maize is equal to
two-fifths of the parental deviation. A cob with 20 rows means a
deviation of 7 from the average of 13, the incipient value of my race.
To reach such an average at once, an ear would be required with 7 x 5/2
= 17-1/2 rows above the average, or an ear with 30-32 rows. These never
occur, but the rule given in a previous lecture gives a method of
calculating the probability of their occurrence, or in other words, the
number of ears required to give a chance of finding such an ear. It
would take too long to give this calculation here, but I find that
approximately 12,000 ears would be required to give one with 28 rows,
which was the highest number attained in [784] my experiment, while
100,000 ears would afford a chance of one with 32 rows*. Had I been able
to secure and inspect this number of ears, perhaps I would have needed
only a year to get an average of 20 rows. This however, not being the
case, I have worked for seven years, but on the other hand have
cultivated all in all only about one thousand individuals for the entire
experiment.


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